The show focuses on the efforts of Dick Dastardly and his canine sidekick Muttley to catch Yankee Doodle Pigeon, a carrier pigeon who carries secret messages (hence the name of the show’s theme song "Stop the Pigeon").
Each story features variations on the same plot elements: the Vulture Squadron tries to trap Yankee Doodle Pigeon using one or more planes equipped with Klunk's latest contraption(s), but one or more of the Squadron messes up and the plane(s) either crash, collide or explode (or even all of the above!). While they are falling out of the wreckage, Dick Dastardly calls for help, which Muttley either offers or refuses depending on whether Dastardly agrees to give him a medal. Even when Muttley does agree to fly Dastardly out of trouble, Dastardly seldom has a soft landing. At some point the General calls Dastardly on the phone to demand results. Dastardly assures him that they will soon capture the pigeon, but the General does not believe him and either bellows down the phone or reaches through it and pulls Dastardly's moustache or nose. Klunk then comes up with a new invention and "explains" it in his own unique way. Dastardly says, "What'd he say? What'd he say?" and Zilly interprets, before attempting to run away. Once Muttley has "persuaded" (usually by biting/attacking him) Zilly to return, the Vulture Squadron take off in their new plane(s) to repeat the whole procedure. Eventually, the Squadron is left to lick their wounds as Yankee Doodle Pigeon flies over the horizon, blowing his bugle triumphantly.
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